ORCHIDS CONSTITUTE the most diverse family of flowering plants with 28,484 species (Govaerts et al., 2017) representing ca. 8% of all vascular plants worldwide (Dressler, 2005; Scotland and Wortley, 2003). The drivers behind this diversification appear to be linked to several factors especially co-evolution with specific insect pollinators needed for sexual reproduction. The majority (60-70%) of orchid species entice and reward pollinators within a blend of attractants that typically include oils, waxes, fragrance (aromatic compounds) and sugar-rich nectar (Ackerman, 1986; Brzosko and Mirski, 2021; Dressler, 1981; Tremblay et al., 2005). About 35 million years ago, orchid speciation accelerated when the family adapted to an arboreal lifestyle aided by their ability to conserve water via crassulacean acid metabolism or CAM (Stokstad, 2015), and flowers targeted winged insects especially bees (Hymenoptera) and moths (Lepidoptera) for pollination. This timeframe coincides with the evolutionary appearance of nocturnal hawk moths (Sphingidae; Kawahara et al., 2019) that pollinate many members of the Epidendroideae, the largest orchid subfamily. This subfamily includes the well-known angraecoids of Africa and Madagascar (Angraecum sesquipedale Thouars; Pridgeon et al., 1999, 2014), and members of the genus Dendrophylax in the New World. Collectively, moth-pollinated orchids are characterized by their white flowers that emit a sweetsmelling evening fragrance, and long nectar spurs that often match the proboscis length of the pollinator (Dressler, 1981; Faegri and van der Pijl, 1979; Pal et al., 2019; Prakash and Pathak, 2020). These spurs are often filled with copious amount of sugar-rich nectar that serves to reward the pollinator with a carbohydrate source needed to power muscles for rapid flight over long distances.